The following is an excerpt of JED Fox's PhD thesis. The article stays fairly true to the original with minor changes. Most measurements are changed to the Metric System.

 

Fox's divided the Lower Montane Forest into 3 main sub-formations:

 

Upper Dipterocarp Forest

This type is not abundant, occupying a transitional position between lower level forests rich in dipterocarp species and higher levels dominated by conifers and Fagaceae. It is largely confined to the eastern slopes of Mount Kinabalu petering out at 1000–1500 m, though individuals or small stands occur on other mountains. The structure and composition of typical stands have not been described, and the full range of species is not known. Nicholson (1962 b) noted Shorea nebulosa at 1000 m in Oak forest on sandstone and shales, and S. platyclados with some S. faguetiana off ridges at 1200m.

 

 Corner (1962 a) noted S. monticola in gregarious stands at 1200–1500 m, generally on peaks, while he observed S. platyclados at 1000 m. In Brunei, Ashton (1964 c) recorded S. monticola in small groups on white sand soil at 1750 m, but also on lower mountains at 900 m. Other observations from Kinabalu area are of S. monticola at the summit of Mt. Hampuan, 1200 m in a matrix of conifers and with S. nebulosa, S. platyclados, Vatica spp., and Hopea spp. at 1500–1800m in the Kundasang/Tenompok area on the southern side of the mountain in a region of conifers and Fagaceae (Meijer 1963 b).

 

Montane species of dipterocarps have been collected in the Crocker Range in the Tambunan/Sinsuron area, on Mt Lumaku (Gunong Lumaku FR), in Mt Templer FR and on Mt Trus Madi. Several species occur from about 700 m in Gunong Rara FR.

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Agathis dammara Forest

This species occurs in gregarious stands at 1600 m near Park Headquarters to the south of Kinabalu, and is locally frequent on the eastern side at 1200–1700 m. A fine stand was seen northeast of Gunung Alab in the Crocker Range at about 1700m. Stands occur in the Serundong Hills of Kalabakan FR and isolated clumps occur in the Tawau mountains, e.g. Mts. Andrassay and Wullersdorf (600–650 m). Tree sizes exceed 60 cm diameter and 24 m in height.

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Oak / Conifer Forest

This is the most extensive form of mountain forest and is present on all the higher parts of the Crocker Range above the zone of shifting cultivation, as well as on the flanks of Kinabalu. Meijer (1963 b) gives a map of Kinabalu based on aerial photography suggesting that this type extends from 900-1500 m on the eastern side and 1500–2100 m on the south. It must be stressed that altitudinal limits, especially the lower, by no means strictly coincide with the contour lines (van Steenis 1962) and the effects of 'mass elevation' are everywhere evident. Shifting cultivation at present extends to 1200–1500 m on the south and west of Kinabalu though primary forest existed at Tenompok around 1910 (Gibbs 1913). The areas south of Park Headquarters, Sosopodon, and north of Kundasang have been cleared off only in recent years. The best development of this type in the Crocker Range is from 1200 to 1300 m.

 

It is not possible to do justice here to the flora of Kinabalu whose wealth of forms and species is enormous: one collector is said to have obtained 200 species of the orchid genus Bulbophyllum (Corner 1962 b). Details of the flora may be found in Meijer (1963 b) and a preliminary account of Pinusok Plateau in Corner (1962 a). Of the conifers large trees of Phyllocladus hypophyllus. Podocarpus imbricatus and Podocarpus neriifolius occur in this type, together with smaller trees of Dacrydium falciforms and Dacrydium beccarii. These are found with many species of Lithocarpus e.g. L. clementianus, L. confertus, L. echinulatus, L. encleisacarpus, L. hallierri, L. lampadarius, L. luteus, L. pappillifer, L. revolutus, L. some Castanopsis e.g. C. acuminatissima and C. clemensii.Other species of Lithocarpus which extend into higher altitudes are L. bullatus, L. havilandii, L. nodosus and L. turbinatus. The type may be described as a two-layered forest in the sense of Robbins (1969) with a thin understorey consisting of species of the families Lauraceae, Theaceae, Clusiaceae, and of Eugenia (Syzygium) species. In admixture with the oaks and conifers, sometimes reaching large sizes, are species of the lowland genera of Moraceae, Meliaceae, Myristicaceae etc.

 

Physiognomy of the mountain forest varies with topography, exposure and soil. In gullies groves of larger trees extend to higher elevations, while on exposed sites trees are smaller. In the Mamut valley, the mossy forest descends to 1500 m due to the effect of low cloud and mist. Similarly hill tops at 1200 m sometimes have low coniferous-myrtaceous forest also found on higher ridges. In general, heights are from 10–30 m, bole sizes reach 60 cm diameter in better growth. Local stands of thin boled Tristaniopsis/Eugenia (Syzygium) occur on white sandy soils (lower montane kerangas?) throughout the type and Corner noted that Gymnostoma sumatrana at 1650 m and riverine stands of Tristaniopsis, Engelhardtia, Duabanga, Adinandra etc. in Mamut area at 1100–1350 m. The (rare) 'trig oak', Trigobalanus verticillata, is locally abundant on steep slopes and ridges at 1200–1500 m.

 

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