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Type
E: Parashorea malaanonan / Dryobalanops Forest
The following is an excerpt of JED Fox's PhD
thesis. The article stays fairly true to the original with minor changes. Most
measurements are changed to the Metric System.
Quick Jump Silabukan
Semporna/Tawau Elsewhere
Words of wisdom
Forests in Silabukan FR*
This Type is primarily found on steeper slopes than
the others described thus far and may be considered as marginal between lowland
and the hill dipterocarp types which follow (i.e. Types F
and G). In eastern
Sabah, this Type is present in
Silabukan FR on mixed volcanic and
sedimentary soils where S. johorensis and Eusideroxylon
zwageri are common associates,
and presence of these is one of the main distinctions between Types A
and E in this area. A relascope survey in Block 42 near Pangaruwon River in
Silabukan FR sampling 70 acres (28 ha) indicates more large trees than
similar surveys in Type A stands in the vicinity.
Stocking data (commercial species only) from six 1 ha
plots in the area felled in 1969 (RP 245/3) in the Bagahak
Range area of Silabukan FR showed that Eusideroxylon
was absent in this rather hilly area. Of particular note are absence of Dipterocarpus species other than D. caudiferus, predominance of
S. johorensis of the Rubroshorea, absence of Selangan Batu
(though Type F is locally represented by S.
laevis within the reserve), and abundance of Richetia mainly S.
hopeifolia
and
S. faguetiana.
This area had representation of smaller trees 1–5
ft girth (10–57 cm diameter) of commercial species totaling 19.7/acre
(48.6/ha) with 27% Richetia, 22 % S.
johorensis, 13% P. malaanonan and
10% Dr. lanceolata. As with other Parashorea
forests, the canopy was irregular and distribution of large trees patchy but
with sizes well over 16 ft girth (166 cm diameter). Of the non-dipterocarps
present, Sympetalandra borneensis was particularly abundant, species of Annonaceae
in the understorey.
*Note: In the early 1970s, Silabukan FR was to a
very large area in the Dent Peninsula. It is
similar to the present areas covering the eastern half of Tabin Wildlife
Reserve and the area southwards to the Lahad Datu--Tungku main road.
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Forests
of the Semporna/Tawau Peninsula
S. johorensis
is common in parts of Kalumpang FR north of Type
D, on the Kalumpang Formation and is also abundant on remaining basalt in
the Quoin area and on andesite, dacite etc. in the Tawau Hills FR with the two
type species. Nicholson (in Paton 1959) reported the same form of forest on Kawa
soils developed on intrusive doleritic rock. Locally very large trees were found
in the Quoin area {e.g. a S. superba SB
of 32 ft girth (310 cm diam)} at 5 m from the ground with total height of 76 ,
but relatively short boles were attributed to the lack of a competing main
storey; blanks with climber tangles were frequent throughout the area of basalt
soils. Mean values for 11 half-acre plots on basalt soils showed total numbers
over 1 ft g (10 cm diam) as 145/acre, with basal area of 170 sq. ft/acre of
which 31 were dipterocarps and 3.4/acre over 8 ft g of which 6.5 were
dipterocarps. Dipterocarpus was
absent, except as small trees. Recent plots established across Quoin (0.45 ha) and
Bald Hills (0.28 ha), both basalt, showed that other dipterocarp species present in these areas but
not included in the samples were S.
macroptera Ru, S. superba SB,
Parashorea
smythiesii, Hopea beccariana and Dipterocarpus
applanatus, the latter as a small tree only.
Tawau Hills FR has long been considered an area of
hill protection forest on highly erodable dacite and andesite hills from
170–300 m altitude. They contained variable stands in alluvial valleys, on
steep slopes and high crests. The latter higher up and outside felling areas
were not sampled but contained forests of Type F.
The P. malaanonan/Dr.
lanceolata Type is represented at lower levels in the mountainous Gunong
Rara FR Plots placed at Miles 39–41 along Luasong/Gunong Rara Road reveal the
absence of Eusideroxylon, scarcity of certain
Rubroshorea species, e.g. S. leprosula and S. ovalis,
scarcity of Dipterocarpus species, increased frequency of Dr. lanceolata, compared with the Luasong area and high
representation of Parashorea. S.
pauciflora, in addition to the other 2 Rubroshorea , is comparatively
abundant Affinities of the forest in this area clearly lie with Type
B to the north in Kuamut FR and to Type C
nearer the coast in Kalabakan FR. Limited occurrence of the Type E
assemblage of species is known for shale ridges in the higher hills of Gunong
Rara, but its full extent is not present known.
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The forests elsewhere
The Type also occurs on the basalt mountains in the Labuk
valley area of central where Meijer (1968) noted high incidence of Pinorea
longiracemosa and Diospyros species.
The same forest probably occurs on the isolated grandiorite peaks northeast of Ranau
in the headwaters of the Sugut River near Paginatan. Relic gallery forest
containing S. acuminatissima (Ri), E.
zwageri and P. malaanonan in the Sook Plains suggests that this type was
formerly present in the area on present day grassland areas of non-podsol soils.
Further west the forests in Mandalom FR is partly of this type and partly
of Type F Selangan Batu Forest. Here Type E is
represented by P. malaanonan and S.
acuminatissima on the lower slopes petering out at above 600 m altitude.
At Pangie on the lower slope of hills in the Gunung
Lumaku FR where Tamburong Formation extends into the Crocker
Range, RP 15 is
representative of the P. malaanonan/Dr. lanceolata Type. In this area it again gives way
to Type F on the steeper higher slopes.
Elsewhere in the Crocker Range, this type has
been largely eliminated (as with Type D) by shifting cultivation, though it
still present inside, and adjacent to Mt. Templar FR. Here Richetia is
represented by S. faguetiana and S.
kudatensis; S. leptoclados is
uncommon and Rubroshorea is represented by
S. ferruginea and S. smithiana;
and both the type species are abundant. Small patches remain in the Sir James
Brook Range at the northern extremity of the Crocker Range, on the lower slopes
of Timbang Batu Hills to the southeast of Marudu Bay, and in Labuan
FR near Kudat. Dipterocarpus warburgii is
present in this forest Type in both Gunong Lumaku between Tenom and
Tomani.
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Words
of wisdom
At present the only important area of commercial
managed forest of Type E is that in Silabukan FR as the type generally occurs on
steeper slopes exploitation is both difficult and potentially more likely to
lead to severe erosion than with Types B and C.
However the principal species components are important in regeneration
considerations where P. malaanonan, S.
johorensis or Dr. lanceolata are
abundant within the main commercial forests.
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